The Cerritos ensigns must assist a caretaker on the voyage of a historically significant starship.

Mariner tries to get demoted, Rutherford tries to get promoted, Boimler makes a big move.

Boimler leads his first away mission on an alien megastructure.

Tendi is summoned back to Orion for a wedding.

The hedonistic outlook of a Betazoid delegation infects the crew.

The Cerritos visits the Ferengi homeworld.

Three computerized villains return to cause problems for the Cerritos crew.

The Lower Deckers go on a classic cave mission.

Captain Freeman assigns the Lower Deckers an overly safe mission to try and keep a self-destructive Mariner out of danger.

Mariner finds herself held as the hoped-for ally of the disgraced Nick Locarno, now a megalomaniacal insurrectionist armed with a rebel fleet and a destructively planet-reforming Genesis device.